Bolitoglossa schizodactyla Wake & Brame, 1966
Accepted species in Catalogue of Life’s 11 September 2026 base release. This preserves the source classification and available supporting data; it is not a completed species monograph.

Bolitoglossa schizodactyla salamander
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Open encyclopedia reference
The Cocle salamander (Bolitoglossa schizodactyla), also known as the Cocle mushroomtongue salamander, is a species of salamander in the family Plethodontidae. It is found in Costa Rica and Panama. In Costa Rica, it is only known from the southeastern part of the country on the Atlantic versant, whereas in Panama it is more widespread and occurs also on the Pacific versant.
Its natural habitats are humid lowland and montane forests. It is a common species in Panama, whereas it is only known from a single specimen in Costa Rica. It is threatened by habitat loss.
Wikipedia contributors, Cocle salamander; see linked history · CC BY-SA 4.0 . Plain-text lead excerpt, at most 180 words; no infobox or later sections Retrieved 2026-10-05T15:27:13.308762+00:00. This reference remains separate from the taxonomic record.
Identity & source status
- Scientific name
- Bolitoglossa schizodactyla Wake & Brame, 1966
- Name authority
- Wake & Brame, 1966
- Rank
- species
- Accepted status
- accepted
- Kingdom
- Animalia
- Phylum
- Chordata
- Class
- Amphibia
- Order
- Caudata
- Family
- Plethodontidae
- Genus
- Bolitoglossa
- Extinction status
- Recorded extant
Classification family tree
Nested taxonomic groups from the source. This is not a direct-ancestor chain, a dated evolutionary tree or a phylogenetic reconstruction.
- domainEukaryota (Chatton, 1925) Whittaker & Margulis, 1978
- kingdomAnimalia
- phylumChordata
- subphylumVertebrata
- infraphylumGnathostomata
- parvphylumOsteichthyes
- megaclassTetrapoda
- classAmphibia
- orderCaudata
- familyPlethodontidae Gray, 1850
- subfamilyHemidactyliinae Hallowell, 1856
- genusBolitoglossa Duméril, Bibron & Duméril, 1854
- speciesBolitoglossa schizodactyla Wake & Brame, 1966
Close classified relatives
These entries share the source’s genus, Bolitoglossa. They are not necessarily the closest relatives established by molecular evidence.
| Species | Classification | Source status |
|---|---|---|
| Bolitoglossa compacta Wake, Brame & Duellman, 1973 | PlethodontidaeChordata | Accepted speciesRecorded extant |
| Bolitoglossa robinsoni Bolaños & Wake, 2009 | PlethodontidaeChordata | Accepted speciesRecorded extant |
| Bolitoglossa splendida Boza-Oviedo, Rovito, Chaves, García-Rodríguez, Artavia, Bolaños & Wake, 2012 | PlethodontidaeChordata | Accepted speciesRecorded extant |
| Bolitoglossa flavimembris (Schmidt, 1936) | PlethodontidaeChordata | Accepted speciesRecorded extant |
| Bolitoglossa alberchi García-París, Parra-Olea, Brame & Wake, 2002 | PlethodontidaeChordata | Accepted speciesRecorded extant |
| Bolitoglossa subpalmata (Boulenger, 1896) | PlethodontidaeChordata | Accepted speciesRecorded extant |
| Bolitoglossa veracrucis Taylor, 1951 | PlethodontidaeChordata | Accepted speciesRecorded extant |
| Bolitoglossa mulleri (Brocchi, 1883) | PlethodontidaeChordata | Accepted speciesRecorded extant |
| Bolitoglossa awajun Cusi, Gagliardi-Urrutia, Brcko, Wake & von May, 2020 | PlethodontidaeChordata | Accepted speciesRecorded extant |
| Bolitoglossa bolanosi Arias, Chaves & Parra-Olea, 2024 | PlethodontidaeChordata | Accepted speciesRecorded extant |
| Bolitoglossa altamazonica (Cope, 1874) | PlethodontidaeChordata | Accepted speciesRecorded extant |
| Bolitoglossa alvaradoi Taylor, 1954 | PlethodontidaeChordata | Accepted speciesRecorded extant |
Source literature & publication evidence
References attached to this source record, including nomenclatural citations where supplied. This is a selected bibliography; abbreviated citations remain in the source’s form.
- https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=208418 Source fields: Taxon.tsv dwc:namePublishedIn, Taxon.tsv dcterms:references
Origins, evidence & remaining research
Name authority and publication references document nomenclature where the source supplies it. They do not establish evolutionary origins or the earliest fossil occurrence.
Specimen-level evidence, morphology, ecological research, phylogenetic studies, a comprehensive bibliography and a located literary passage have not yet been compiled for this record.
Dataset citation & attribution
This exact taxon in ChecklistBank · Catalogue of Life 2026-09-11
Catalogue of Life 2026-09-11; dataset 316321; doi:10.48580/dgz5n. https://creativecommons.org/licenses/by/4.0/ . Retrieved 5 October 2026.
Stable key: col:MDS3. Classification follows source parent identifiers. English common names may vary by region.