Tomopterna tuberculosa (Boulenger, 1882)
Accepted species in Catalogue of Life’s 11 September 2026 base release. This preserves the source classification and available supporting data; it is not a completed species monograph.
Open encyclopedia reference
The rough sand frog (Tomopterna tuberculosa) is a species of frog in the family Pyxicephalidae.
It is found in Angola, the Democratic Republic of the Congo, Namibia, Tanzania, Zambia, and Zimbabwe, and possibly Malawi and Mozambique.
Its natural habitats are dry savanna, moist savanna, freshwater marshes, and intermittent freshwater marshes.
Wikipedia contributors, Rough sand frog; see linked history · CC BY-SA 4.0 . Plain-text lead excerpt, at most 180 words; no infobox or later sections Retrieved 2026-10-05T18:54:05.697441+00:00. This reference remains separate from the taxonomic record.
Identity & source status
- Scientific name
- Tomopterna tuberculosa (Boulenger, 1882)
- Name authority
- (Boulenger, 1882)
- Rank
- species
- Accepted status
- accepted
- Kingdom
- Animalia
- Phylum
- Chordata
- Class
- Amphibia
- Order
- Anura
- Family
- Pyxicephalidae
- Genus
- Tomopterna
- Extinction status
- Recorded extant
Classification family tree
Nested taxonomic groups from the source. This is not a direct-ancestor chain, a dated evolutionary tree or a phylogenetic reconstruction.
- domainEukaryota (Chatton, 1925) Whittaker & Margulis, 1978
- kingdomAnimalia
- phylumChordata
- subphylumVertebrata
- infraphylumGnathostomata
- parvphylumOsteichthyes
- megaclassTetrapoda
- classAmphibia
- orderAnura
- familyPyxicephalidae Bonaparte, 1850
- subfamilyCacosterninae Noble, 1931
- genusTomopterna Duméril & Bibron, 1841
- speciesTomopterna tuberculosa (Boulenger, 1882)
Close classified relatives
These entries share the source’s genus, Tomopterna. They are not necessarily the closest relatives established by molecular evidence.
| Species | Classification | Source status |
|---|---|---|
| Tomopterna delalandii (Tschudi, 1838) | PyxicephalidaeChordata | Accepted speciesRecorded extant |
| Tomopterna gallmanni Wasonga & Channing, 2013 | PyxicephalidaeChordata | Accepted speciesRecorded extant |
| Tomopterna kachowskii (Nikolskii, 1900) | PyxicephalidaeChordata | Accepted speciesRecorded extant |
| Tomopterna wambensis Wasonga & Channing, 2013 | PyxicephalidaeChordata | Accepted speciesRecorded extant |
| Tomopterna ahli (Deckert, 1938) | PyxicephalidaeChordata | Accepted speciesRecorded extant |
| Tomopterna pulchra (Boulenger, 1896) | PyxicephalidaeChordata | Accepted speciesRecorded extant |
| Tomopterna branchi Wilson & Channing, 2019 | PyxicephalidaeChordata | Accepted speciesRecorded extant |
| Tomopterna krugerensis Passmore & Carruthers, 1975 | PyxicephalidaeChordata | Accepted speciesRecorded extant |
| Tomopterna luganga Channing, Moyer & Dawood, 2004 | PyxicephalidaeChordata | Accepted speciesRecorded extant |
| Tomopterna marmorata (Peters, 1854) | PyxicephalidaeChordata | Accepted speciesRecorded extant |
| Tomopterna milletihorsini (Angel, 1922) | PyxicephalidaeChordata | Accepted speciesRecorded extant |
| Tomopterna monticola (Fischer, 1884) | PyxicephalidaeChordata | Accepted speciesRecorded extant |
Names & synonyms
Names explicitly linked to this accepted taxon in the source. Synonymy can change with revision.
| Recorded name | Source status |
|---|---|
| Pyxicephalus tuberculosus (Boulenger, 1882) (Boulenger, 1882) | synonym |
| Rana cacondana Ahl, 1927 Ahl, 1927 | synonym |
| Rana signata Ahl, 1927 Ahl, 1927 | synonym |
| Pyxicephalus rugosus Günther, 1865 Günther, 1865 | synonym |
| Tomopterna rugosa (Günther, 1865) (Günther, 1865) | synonym |
| Rana tuberculosa Boulenger, 1882 Boulenger, 1882 | synonym |
Source literature & publication evidence
References attached to this source record, including nomenclatural citations where supplied. This is a selected bibliography; abbreviated citations remain in the source’s form.
- https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=665475 Source fields: Taxon.tsv dwc:namePublishedIn, Taxon.tsv dcterms:references
Origins, evidence & remaining research
Name authority and publication references document nomenclature where the source supplies it. They do not establish evolutionary origins or the earliest fossil occurrence.
Specimen-level evidence, morphology, ecological research, phylogenetic studies, a comprehensive bibliography and a located literary passage have not yet been compiled for this record. A suitable sourced image remains to be added.
Dataset citation & attribution
This exact taxon in ChecklistBank · Catalogue of Life 2026-09-11
Catalogue of Life 2026-09-11; dataset 316321; doi:10.48580/dgz5n. https://creativecommons.org/licenses/by/4.0/ . Retrieved 5 October 2026.
Stable key: col:57DS5. Classification follows source parent identifiers. English common names may vary by region.