Okinawa Island Spiny Rat
Tokudaia muenninki (Johnson, 1946)
Accepted species in Catalogue of Life’s 11 September 2026 base release. This preserves the source classification and available supporting data; it is not a completed species monograph.

geographic distribution of Tokudaia muenninki
IUCN. Assessor/s: Ishii, N. & Kaneko, Y. Reviewer/s: Amori, G. (Small Nonvolant Mammal Red List Authority) & Chanson, J. (Global Mammal Assessment Team) · CC BY-SA 3.0 · Image sourceProvider-generated thumbnail requested at 800 pixels; no local editing Local WebP display rendition; resized without cropping.
Open encyclopedia reference
Muennink's spiny rat or Okinawa spiny rat (Tokudaia muenninki, Japanese: オキナワトゲネズミ, romanized: Okinawa togenezumi or 沖縄棘鼠) is a species of rodent in the family Muridae. Endemic to Okinawa Island, Japan, its natural habitat is subtropical moist broadleaf forest. The karyotype has 2n = 44. Its sex chromosomes are abnormally large, while the other two species in Tokudaia have lost their Y chromosome. It is found only on the northern part (Yanbaru area) of the island, above 300 m, and is thought to inhabit an area of less than 3 km2. The head and body are up to 7 inches long with a 5-inch tail. They weigh up to 7 ounces. They have a short thick body and dense fur, consisting of fine hairs and coarse, grooved spines (hence the common name "spiny rat"). The fur is brownish above and grayish white below with a faint orange tinge. The spines on the animal's back are black throughout while the spines underneath are usually white with a reddish-brown tip.
Wikipedia contributors, Muennink's spiny rat; see linked history · CC BY-SA 4.0 . Plain-text lead excerpt, at most 180 words; no infobox or later sections Retrieved 2026-10-05T12:39:53.570669+00:00. This reference remains separate from the taxonomic record.
Identity & source status
- Scientific name
- Tokudaia muenninki (Johnson, 1946)
- Name authority
- (Johnson, 1946)
- Common name
- Okinawa Island Spiny Rat
- Rank
- species
- Accepted status
- accepted
- Kingdom
- Animalia
- Phylum
- Chordata
- Class
- Mammalia
- Order
- Rodentia
- Family
- Muridae
- Genus
- Tokudaia
- Extinction status
- Recorded extant
Classification family tree
Nested taxonomic groups from the source. This is not a direct-ancestor chain, a dated evolutionary tree or a phylogenetic reconstruction.
- domainEukaryota (Chatton, 1925) Whittaker & Margulis, 1978
- kingdomAnimalia
- phylumChordata
- subphylumVertebrata
- infraphylumGnathostomata
- parvphylumOsteichthyes
- megaclassTetrapoda
- classMammalia Linnaeus, 1758
- subclassTheria Parker & Haswell, 1897
- infraclassEutheria Gill, 1872
- orderRodentia Bowdich, 1821
- suborderMyomorpha Brandt, 1855
- superfamilyMuroidea Illiger, 1811
- familyMuridae Illiger, 1811
- subfamilyMurinae Illiger, 1811
- genusTokudaia Kuroda, 1943
- speciesTokudaia muenninki (Johnson, 1946)
Close classified relatives
No other member of this genus is present in the selected snapshot.
Documented distribution
Source distribution statements, not a complete range map. A listed locality is not necessarily a place of origin.
Source literature & publication evidence
References attached to this source record, including nomenclatural citations where supplied. This is a selected bibliography; abbreviated citations remain in the source’s form.
- https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=585600 Source fields: Taxon.tsv dwc:namePublishedIn, Taxon.tsv dcterms:references
Origins, evidence & remaining research
Name authority and publication references document nomenclature where the source supplies it. They do not establish evolutionary origins or the earliest fossil occurrence.
Specimen-level evidence, morphology, ecological research, phylogenetic studies, a comprehensive bibliography and a located literary passage have not yet been compiled for this record.
Dataset citation & attribution
This exact taxon in ChecklistBank · Catalogue of Life 2026-09-11
Catalogue of Life 2026-09-11; dataset 316321; doi:10.48580/dgz5n. https://creativecommons.org/licenses/by/4.0/ . Retrieved 5 October 2026.
Stable key: col:57BC9. Classification follows source parent identifiers. English common names may vary by region.