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honeycomb stingray

Himantura uarnak (Gmelin, 1789)

Source-backed taxonomic record

Accepted species in Catalogue of Life’s 11 September 2026 base release. This preserves the source classification and available supporting data; it is not a completed species monograph.

A fine-spotted leopard whipray (Himantura tutul) at the California Academy of Sciences' Philippine Mangrove Lagoon exhibit.

A fine-spotted leopard whipray (Himantura tutul) at the California Academy of Sciences' Philippine Mangrove Lagoon exhibit.

User:Steven Walling · CC BY-SA 3.0 · Image source

Provider-generated thumbnail requested at 800 pixels; no local editing Local WebP display rendition; resized without cropping.

Open encyclopedia reference

The reticulate whipray or honeycomb stingray (Himantura uarnak) is a species of stingray in the family Dasyatidae. It inhabits coastal waters in the western Indian Ocean including the Red Sea, Natal and the Arabian Sea; also a Lessepsian transmigrant in the eastern Mediterranean. A large species reaching 2 m (6.6 ft) in width, the reticulate whipray has a diamond-shaped pectoral fin disc and an extremely long tail without fin folds. Both its common and scientific names refer to its ornate dorsal color pattern of many small, close-set dark spots or reticulations on a lighter background. However, the reticulate whipray is only one of several large spotted stingrays in the Indo-Pacific which, coupled with the variability of its coloration with age and locality, has resulted in a great deal of taxonomic confusion. Often encountered resting on the bottom during daytime, the reticulate whipray is a predator of bottom-dwelling invertebrates and bony fishes. Like other stingrays, it is aplacental viviparous, with the developing embryos nourished at first by yolk and later by histotroph ("uterine milk").

Wikipedia contributors, Reticulate whipray; see linked history · CC BY-SA 4.0 . Plain-text lead excerpt, at most 180 words; no infobox or later sections Retrieved 2026-10-05T07:26:54.466025+00:00. This reference remains separate from the taxonomic record.

Identity & source status

Scientific name
Himantura uarnak (Gmelin, 1789)
Name authority
(Gmelin, 1789)
Common name
honeycomb stingray
Rank
species
Accepted status
accepted
Kingdom
Animalia
Phylum
Chordata
Class
Elasmobranchii
Order
Myliobatiformes
Family
Dasyatidae
Genus
Himantura
Extinction status
Not supplied by this source

Source habitat flags: Marine: yes · Freshwater: no · Terrestrial: no. These do not form a complete habitat description.

Classification family tree

Nested taxonomic groups from the source. This is not a direct-ancestor chain, a dated evolutionary tree or a phylogenetic reconstruction.

  1. domainEukaryota (Chatton, 1925) Whittaker & Margulis, 1978
  2. kingdomAnimalia
  3. phylumChordata
  4. subphylumVertebrata
  5. infraphylumGnathostomata
  6. parvphylumChondrichthyes
  7. classElasmobranchii
  8. subclassNeoselachii
  9. infraclassBatoidea
  10. orderMyliobatiformes
  11. familyDasyatidae Jordan & Gilbert, 1879
  12. genusHimantura Müller & Henle, 1837
  13. speciesHimantura uarnak (Gmelin, 1789)

Close classified relatives

These entries share the source’s genus, Himantura. They are not necessarily the closest relatives established by molecular evidence.

SpeciesClassificationSource status
Himantura australis Last, White & Naylor, 2016DasyatidaeChordataAccepted speciesExtinction status not supplied
Himantura krempfi (Chabanaud, 1923)DasyatidaeChordataAccepted speciesExtinction status not supplied
Himantura marginata (Blyth, 1860)DasyatidaeChordataAccepted speciesExtinction status not supplied
Himantura pareh (Bleeker, 1852)DasyatidaeChordataAccepted speciesExtinction status not supplied
Himantura arlyzae Borsa, Hernawan, Hocdé & Pouyaud, 2026DasyatidaeChordataAccepted speciesExtinction status not supplied
Himantura alcockii (Annandale, 1909)DasyatidaeChordataAccepted speciesExtinction status not supplied
Himantura fava (Annandale, 1909)DasyatidaeChordataAccepted speciesExtinction status not supplied
Himantura fluviatilis (Hamilton, 1822)DasyatidaeChordataAccepted speciesExtinction status not supplied
Himantura leoparda Manjaji-Matsumoto & Last, 2008DasyatidaeChordataAccepted speciesExtinction status not supplied
Himantura microphthalma (Chen, 1948)DasyatidaeChordataAccepted speciesExtinction status not supplied
Himantura tutul Borsa, Durand, Shen, Alyza, Solihin & Berrebi, 2013DasyatidaeChordataAccepted speciesExtinction status not supplied
Himantura undulata (Bleeker, 1852)DasyatidaeChordataAccepted speciesExtinction status not supplied

Browse 85 records assigned to Dasyatidae.

Names & synonyms

Names explicitly linked to this accepted taxon in the source. Synonymy can change with revision.

Recorded nameSource status
Dasybatus uarnak (Gmelin, 1789) (Gmelin, 1789)synonym
Himantura punctata (Günther, 1870) (Günther, 1870)synonym
Raja sephen var. uarnak Forsskål, 1775 Forsskål, 1775synonym
Raja uarnak Gmelin, 1789 Gmelin, 1789synonym
Raja sephen uarnak Forsskål, 1775 Forsskål, 1775synonym
Dasyatis uarnak (Gmelin, 1789) (Gmelin, 1789)synonym
Trygon uarnak (Gmelin, 1789) (Gmelin, 1789)synonym
Trygon punctata Günther, 1870 Günther, 1870synonym

Source literature & publication evidence

References attached to this source record, including nomenclatural citations where supplied. This is a selected bibliography; abbreviated citations remain in the source’s form.

  1. https://www.marinespecies.org/aphia.php?p=taxdetails&id=105854 Source fields: Taxon.tsv dwc:namePublishedIn, Taxon.tsv dcterms:references

Origins, evidence & remaining research

Name authority and publication references document nomenclature where the source supplies it. They do not establish evolutionary origins or the earliest fossil occurrence.

Specimen-level evidence, morphology, ecological research, phylogenetic studies, a comprehensive bibliography and a located literary passage have not yet been compiled for this record.

Dataset citation & attribution

This exact taxon in ChecklistBank · Catalogue of Life 2026-09-11

Catalogue of Life 2026-09-11; dataset 316321; doi:10.48580/dgz5n. https://creativecommons.org/licenses/by/4.0/ . Retrieved 5 October 2026.

Stable key: col:3LY4N. Classification follows source parent identifiers. English common names may vary by region.